Touzot, M., Holmstrup, M., Sørensen, J. G. & Slotsbo, S. (2025). Gain of thermal tolerance through acclimation is quicker than the loss by de-acclimation in the freeze-tolerant potworm, Enchytraeus albidus. Journal of Experimental Biology, 228(5), Artikel jeb249675. https://doi.org/10.1242/jeb.249675

Dai, W., Amstrup, A. B., Slotsbo, S., Holmstrup, M., Touzot, M. & Sørensen, J. G. (2025). Increased amplitude of daily temperature fluctuation magnifies the acute toxicity of phenanthrene and upregulates molecular mechanisms of stress defense in Enchytraeus albidus (Oligochaeta). Ecotoxicology and Environmental Safety, 303, Artikel 118802. https://doi.org/10.1016/j.ecoenv.2025.118802

Gerlich, H. S., Holmstrup, M., Schmidt, N. M., Phillimore, A. B. & Høye, T. T. (2025). Keeping up with climate change: have Arctic arthropods reached their phenological limits? Proceedings of the Royal Society B: Biological Sciences, 292(2051), Artikel 20250350. https://doi.org/10.1098/rspb.2025.0350

Gerlich, H. S., Schøn, M. L., Grantland, D., Slotsbo, S. & Holmstrup, M. (2025). Optimizing Enchytraeus albidus Production With Agri-Food Waste: A Nutritional Landscape Approach. Aquaculture Research, 2025(1), Artikel 5632761. https://doi.org/10.1155/are/5632761

Gerlich, H. S., Holmstrup, M., Schmidt, N. M. & Høye, T. T. (2025). Phenological responses to climate change across taxa and local habitats in a high-Arctic arthropod community. Ecological Monographs, 95(1), Artikel e1643. https://doi.org/10.1002/ecm.1643

Wehrli, M., Ge, J., Slotsbo, S. & Holmstrup, M. (2025). Pre-Exposure to Chemicals Increases Springtail Vulnerability to High Temperatures. Global change biology, 31(7), Artikel e70374. https://doi.org/10.1111/gcb.70374

Wang, Y., Slotsbo, S., Bak, S. Y., Martyniuk, C. J. & Holmstrup, M. (2025). Proteomic Responses of the Springtail Folsomia candida to Drought. Insects, 16(7), Artikel 707. https://doi.org/10.3390/insects16070707

Gerlich, H. S., Loboda, S., Simpson, G. L., Savage, J., Schmidt, N. M., Holmstrup, M. & Høye, T. T. (2025). Species' traits modulate rapid changes in flight time in high-Arctic muscid flies under climate change. Proceedings of the Royal Society B: Biological Sciences, 292(2050), Artikel 20250970. https://doi.org/10.1098/rspb.2025.0970

Grantland, D., Holmstrup, M., Overgaard, J., Holmstrup, M. E. & Slotsbo, S. (2025). Tiny Worms, Big Potential: Enchytraeus albidus (Annelida: Clitellata) as Starter Feed for Rainbow Trout. Aquaculture Research, 2025(1), Artikel 7339644. https://doi.org/10.1155/are/7339644

Wehrli, M., Slotsbo, S., Ge, J. & Holmstrup, M. (2024). Acclimation temperature influences the thermal sensitivity of injury accumulation in Folsomia candida at extreme low and high temperatures. Current Research in Insect Science, 6, Artikel 100089. https://doi.org/10.1016/j.cris.2024.100089

Wehrli, M., Slotsbo, S., Fomsgaard, IS., Laursen, BB., Gröning, J., Liess, M. & Holmstrup, M. (2024). A Dirt(y) World in a Changing Climate: Importance of Heat Stress in the Risk Assessment of Pesticides for Soil Arthropods. Global change biology, 30(10), Artikel e17542. https://doi.org/10.1111/gcb.17542

Xie, L., Slotsbo, S., Ilyaskina, D., Forbes, V. & Holmstrup, M. (2024). Chronic Effects of an Insect Growth Regulator (teflubenzuron) on the Life Cycle and Population Growth Rate of Folsomia candida. Environmental Toxicology and Chemistry, 43(5), 1173-1183. https://doi.org/10.1002/etc.5850

Ge, J., Slotsbo, S., Sørensen, J. G. & Holmstrup, M. (2024). Does copper contamination change thermotaxis of the soil arthropod Folsomia candida (Collembola)? Journal of Thermal Biology, 124, Artikel 103950. https://doi.org/10.1016/j.jtherbio.2024.103950

Barezzi, E., Slotsbo, S. & Holmstrup, M. (2024). Does pre-exposure to mild drought enhance desiccation resistance in Collembola? European Journal of Entomology, 121, 303-309. https://doi.org/10.14411/eje.2024.032

Xie, L., Slotsbo, S., Damgaard, C. & Holmstrup, M. (2024). Exposure to teflubenzuron reduces drought tolerance of collembolans. Chemosphere, 361, Artikel 142448. https://doi.org/10.1016/j.chemosphere.2024.142448

Potapov, A. M., Chen, T. W., Striuchkova, A. V., Alatalo, J. M., Alexandre, D., Arbea, J., Ashton, T., Ashwood, F., Babenko, A. B., Bandyopadhyaya, I., Baretta, C. R. D. M., Baretta, D., Barnes, A. D., Bellini, B. C., Bendjaballah, M., Berg, M. P., Bernava, V., Bokhorst, S., Bokova, A. I. ... Scheu, S. (2024). Global fine-resolution data on springtail abundance and community structure. Scientific Data, 11(1), Artikel 22. https://doi.org/10.1038/s41597-023-02784-x, https://doi.org/10.1038/s41597-023-02784-x

Wang, Y., Slotsbo, S., Sørensen, P. B. & Holmstrup, M. (2024). Low soil Moisture Slows Uptake and Elimination Rates of Phenanthrene in Springtails. Bulletin of Environmental Contamination and Toxicology, 112(2), Artikel 25. https://doi.org/10.1007/s00128-024-03854-5

Holmstrup, M., Touzot, M. & Slotsbo, S. (2023). Characterization of the thermal death time landscape for Enchytraeus albidus. Pedobiologia, 97-98, Artikel 150876. https://doi.org/10.1016/j.pedobi.2023.150876

Daly, E. Z., Gerlich, H. S., Frenot, Y., Høye, T. T., Holmstrup, M. & Renault, D. (2023). Climate Change Helps Polar Invasives Establish and Flourish: Evidence from Long-Term Monitoring of the Blowfly Calliphora vicina. Biology, 12(1), Artikel 111. https://doi.org/10.3390/biology12010111

Ge, J., Slotsbo, S., Sørensen, J. G. & Holmstrup, M. (2023). Copper-contaminated soil compromises thermal performance in the springtail Folsomia candida (Collembola). Science of the Total Environment, 897, Artikel 165334. https://doi.org/10.1016/j.scitotenv.2023.165334

Dai, W., Slotsbo, S., Holmstrup, M. & van Gestel, C. A. M. (2023). Evaluation of life-history traits in Folsomia candida exposed to combined repeated mild heat shocks with phenanthrene. Environmental Science and Pollution Research, 30(19), 55132–55142 . https://doi.org/10.1007/s11356-023-26230-y

Potapov, A. M., Guerra, C. A., van den Hoogen, J., Babenko, A., Bellini, B. C., Berg, M. P., Chown, S. L., Deharveng, L., Kováč, Ľ., Kuznetsova, N. A., Ponge, J. F., Potapov, M. B., Russell, D. J., Alexandre, D., Alatalo, J. M., Arbea, J. I., Bandyopadhyaya, I., Bernava, V., Bokhorst, S. ... Scheu, S. (2023). Globally invariant metabolism but density-diversity mismatch in springtails. Nature Communications, 14(1), Artikel 674. https://doi.org/10.1038/s41467-023-36216-6

Dai, W., Holmstrup, M., Slotsbo, S., Bakker, R., Damgaard, C. & van Gestel, C. A. M. (2023). Heat stress delays detoxification of phenanthrene in the springtail Folsomia candida. Chemosphere, 311(Part 2), Artikel 137119. https://doi.org/10.1016/j.chemosphere.2022.137119

Dai, W., Slotsbo, S., Xie, L., Wang, Y., Damgaard, C. & Holmstrup, M. (2023). Increased daily temperature fluctuations exacerbate the toxicity of phenanthrene in Enchytraeus albidus (Enchytraeidae). Science of the Total Environment, 873, Artikel 162403. https://doi.org/10.1016/j.scitotenv.2023.162403

Wang, Y., Slotsbo, S., Damgaard, C. & Holmstrup, M. (2023). Influence of Soil Moisture on Bioaccumulation, Growth, and Recruitment of Folsomia candida Exposed to Phenanthrene-Polluted Soil. Environmental Science & Technology, 57(8), 3085–3094. https://doi.org/10.1021/acs.est.2c07497

Scholl, C. L., Holmstrup, M., Graham, L. A. & Davies, P. L. (2023). Polyproline type II helical antifreeze proteins are widespread in Collembola and likely originated over 400 million years ago in the Ordovician Period. Scientific Reports, 13(1), Artikel 8880. https://doi.org/10.1038/s41598-023-35983-y

Xie, L., Slotsbo, S. & Holmstrup, M. (2023). Tolerance of high temperature and associated effects on reproduction in euedaphic Collembola. Journal of Thermal Biology, 113, Artikel 103439. https://doi.org/10.1016/j.jtherbio.2022.103439

Thakur, M. P., Sigurðsson, B. D., Sigurðsson, P. & Holmstrup, M. (2023). Warming shifts the biomass distribution of soil microarthropod communities. Soil Biology and Biochemistry, 177, Artikel 108894. https://doi.org/10.1016/j.soilbio.2022.108894

Holmstrup, M., Sørensen, J. G., Dai, W., Krogh, P. H., Schmelz, R. M. & Slotsbo, S. (2022). Analysis of heat and cold tolerance of a freeze-tolerant soil invertebrate distributed from temperate to Arctic regions: evidence of selection for extreme cold tolerance. Journal of Comparative Physiology B: Biochemical, Systemic, and Environmental Physiology, 192(3-4), 435-445. https://doi.org/10.1007/s00360-022-01433-w

Holmstrup, M. E. D., Gadeberg, S. F., Engell-Sørensen, K., Slotsbo, S. & Holmstrup, M. (2022). A new strategy in rearing of European flounder: using live Enchytraeus albidus to enhance juvenile growth. Journal of Insects as Food and Feed, 8(11), 1333-1341. https://doi.org/10.3920/JIFF2021.0106

Bonfanti, J., Hedde, M., Cortet, J., Krogh, P. H., Larsen, K. S. & Holmstrup, M. (2022). Communities of Collembola show functional resilience in a long-term field experiment simulating climate change. Pedobiologia, 90, Artikel 150789. https://doi.org/10.1016/j.pedobi.2022.150789

Peng, Y., Holmstrup, M., Schmidt, I. K., De Schrijver, A., Schelfhout, S., Heděnec, P., Zheng, H., Bachega, L. R., Yue, K. & Vesterdal, L. (2022). Litter quality, mycorrhizal association, and soil properties regulate effects of tree species on the soil fauna community. Geoderma, 407, Artikel 115570. https://doi.org/10.1016/j.geoderma.2021.115570

Wang, Y., Slotsbo, S. & Holmstrup, M. (2022). Soil dwelling springtails are resilient to extreme drought in soil, but their reproduction is highly sensitive to small decreases in soil water potential. Geoderma, 421, Artikel 115913. https://doi.org/10.1016/j.geoderma.2022.115913

Jensen, K., Toft, S., Sørensen, J. G. & Holmstrup, M. (2022). Survival and predation rate of wild-caught and commercially produced Orius majusculus (Reuter) (Hemiptera: Anthocoridae). Bulletin of Entomological Research, 112(3), 311-317. https://doi.org/10.1017/S0007485321000055

Marty, A., Boeriis, T., Martínez-De León, G., Holmstrup, M. & Thakur, M. P. (2022). Temperature-dependent trade-offs in maternal investments: An experimental test with two closely related soil microarthropods. European Journal of Soil Biology, 110, Artikel 103402. https://doi.org/10.1016/j.ejsobi.2022.103402

Peng, Y., Holmstrup, M., Kappel Schmidt, I., Ruggiero Bachega, L., Schelfhout, S., Zheng, H., Heděnec, P., Yue, K. & Vesterdal, L. (2022). Tree species identity is the predominant modulator of the effects of soil fauna on leaf litter decomposition. Forest Ecology and Management, 520, Artikel 120396. https://doi.org/10.1016/j.foreco.2022.120396

Krogh, P. H., Lamandé, M., Holmstrup, M. & Eriksen, J. (2021). Earthworm burrow number and vertical distribution are affected by the crop sequence of a grass-clover rotation system. European Journal of Soil Biology, 103, Artikel 103294. https://doi.org/10.1016/j.ejsobi.2021.103294

Phillips, H. R. P., Bach, E. M., Bartz, M. L. C., Bennett, J. M., Beugnon, R., Briones, M. J. I., Brown, G. G., Ferlian, O., Gongalsky, K. B., Guerra, C. A., König-Ries, B., Krebs, J. J., Orgiazzi, A., Ramirez, K. S., Russell, D. J., Schwarz, B., Wall, D. H., Brose, U., Decaëns, T. ... Eisenhauer, N. (2021). Global data on earthworm abundance, biomass, diversity and corresponding environmental properties. Scientific Data, 8, Artikel 136. https://doi.org/10.1038/s41597-021-00912-z

Forbes, V. E., Agatz, A., Ashauer, R., Butt, K. R., Capowiez, Y., Duquesne, S., Ernst, G., Focks, A., Gergs, A., Hodson, M. E., Holmstrup, M., Johnston, A. S. A., Meli, M., Nickisch, D., Pieper, S., Rakel, K. J., Reed, M., Roembke, J., Schäfer, R. B. ... Roeben, V. (2021). Mechanistic Effect Modeling of Earthworms in the Context of Pesticide Risk Assessment: Synthesis of the FORESEE Workshop. Integrated Environmental Assessment and Management, 17(2), 352-363. https://doi.org/10.1002/ieam.4338

Dai, W., Slotsbo, S., van Gestel, C. A. M. & Holmstrup, M. (2021). Temperature-Dependent Toxicokinetics of Phenanthrene in Enchytraeus albidus (Oligochaeta). Environmental Science & Technology, 55(3), 1876-1884. https://doi.org/10.1021/acs.est.0c06182

Bahrndorff, S., Alemu, T., Kristensen, T. N., Sørensen, M. H., Høye, T. T. & Holmstrup, M. (2021). Thermal adaptations of adults and eggs in the Arctic seed bug Nysius groenlandicus (Insecta: Hemiptera) from South Greenland. Polar Biology, 44(3), 491-498. https://doi.org/10.1007/s00300-021-02807-6

Dai, W., Slotsbo, S. & Holmstrup, M. (2021). Thermal optimum for mass production of the live feed organism Enchytraeus albidus. Journal of Thermal Biology, 97, Artikel 102865. https://doi.org/10.1016/j.jtherbio.2021.102865

Engell Dahl, J., Marti, S. L., Colinet, H., Wiegand, C., Holmstrup, M. & Renault, D. (2021). Thermal plasticity and sensitivity to insecticides in populations of an invasive beetle: Cyfluthrin increases vulnerability to extreme temperature. Chemosphere, 274, Artikel 129905. https://doi.org/10.1016/j.chemosphere.2021.129905

Graham, L. A., Boddington, M. E., Holmstrup, M. & Davies, P. L. (2020). Antifreeze protein complements cryoprotective dehydration in the freeze-avoiding springtail Megaphorura arctica. Scientific Reports, 10, Artikel 3047. https://doi.org/10.1038/s41598-020-60060-z

Dai, W., Holmstrup, M., Slotsbo, S., Ke, X., Li, Z., Gao, M. & Wu, L. (2020). Compartmentation and effects of lead (Pb) in the collembolan, Folsomia candida. Environmental Science and Pollution Research, 27(35), 43638–43645. https://doi.org/10.1007/s11356-020-10300-6

Jensen, T. G., Holmstrup, M., Madsen, R. B., Glasius, M., Trac, L. N., Mayer, P., Ehlers, B. & Slotsbo, S. (2020). Effects of α-pinene on life history traits and stress tolerance in the springtail Folsomia candida. Comparative Biochemistry and Physiology Part - C: Toxicology and Pharmacology, 229, Artikel 108681. https://doi.org/10.1016/j.cbpc.2019.108681

Toft, S., Jensen, K., Sørensen, J. G., Sigsgaard, L. & Holmstrup, M. (2020). Food quality of Ephestia eggs, the aphid Rhopalosiphum padi and mixed diet for Orius majusculus. Journal of Applied Entomology (Print), 144(4), 251-262. https://doi.org/10.1111/jen.12739

Gerlich, H. S., Holmstrup, M., Bjerregaard, P. & Slotsbo, S. (2020). Mercury (Hg²⁺) interferes with physiological adaptations to freezing in the arctic earthworm Enchytraeus albidus. Ecotoxicology and Environmental Safety, 204, Artikel 111005. https://doi.org/10.1016/j.ecoenv.2020.111005

Ehlers, B., Berg, M. P., Staudt, M., Holmstrup, M., Glasius, M., Ellers, J., Tomiolo, S., Madsen, R. B., Slotsbo, S. & Peñuelas, J. (2020). Plant Secondary Compounds in Soil and Their Role in Belowground Species Interactions. Trends in Ecology & Evolution, 35(8), 716-730. https://doi.org/10.1016/j.tree.2020.04.001